The Lee Segel Prizes were established in memory of Lee Segel, who made great contributions to the Bulletin of Mathematical Biology and the field of mathematical biology as a whole. The prizes honor outstanding contributions to the field of mathematical biology and will help to promote and advance important research findings in this scientific area. There is a Best Paper Prize, as well as a Best Student Paper Prize. Other prizes may be awarded as deemed appropriate by the Awards Committee, Editors-in-Chief of the Bulletin of Mathematical Biology, and the Society for Mathematical Biology.

The Lee Segel Prizes are awarded every two years, starting in 2009. Papers are selected taking into account the advice of referees and referee reports. For a paper to qualify as a Best Student Paper, the main author of the paper must have been a student at the time that the work was carried out.

The Best Paper Award consists of $3000 USD and an invitation for one of the authors to present the paper at the Annual Meeting of the Society for Mathematical Biology (preferably in the year that the award is given out). The Best Student Paper Award consists of $2000 USD and an invitation for one of the authors to present the paper at the Annual Meeting of the Society for Mathematical Biology.

**Recipients of the Lee A. Segel Prize**

**2021**

**Best Paper: **C. A. Yates, M. J. Ford and R. L. Mort, A multi-stage representation of cell proliferation as a Markov process. *Bull Math Biol* (2017) **79b** 2905-2928. DOI: 10.1007/s11538-013-9827-4.

**Best Student Paper: **A. P. Browning, P. Haridas and M. J. Simpson, A Bayesian sequential learning framework to parameterise continuum models of melanoma invasion into human skin. *Bull Math Biol *(2019) **81**, 676-698. DOI: 10.1007/s11538-018-0532-1.

**2019**

**Best Paper: **M. P. Thon, H. Z. Ford, M. W. Gee and M. R. Myerscough, a quantitative model of atherosclerotic plaques using *in vitro* experiments. *Bull Math Biol *(2017) **80**, 175-214. DOI: 10.1007/s11538-013-9827-4.

**Best Student Paper: **M. Craig, A. R. Humphries and M. C. Mackay, A mathematical model of granulopoiesis incorporating the negative feedback dynamics and kinetics of G-CSF/neutrophil binding and internalisation. *Bull Math Biol *(2016) **78**, 2304-2357. DOI: 10.1007/s11538-013-9827-4.

**2017**

**Best Paper: **A. J. McKane, T. Biancalani and T. Rogers. Stochastic pattern formation and spontaneous polarisation: The linear noise approximation and beyond.* Bull Math Biol *(2014) **76, **895-921. DOI: 10.1007/s11538-013-9827-4.

**Best Student Paper (shared prize): **J. P. Taylor-King, E. E. van Loon, G. Rosser and S. J. Chapman. From birds to bacteria: generalised velocity jump processes with resting states.* Bull Math Biol* (2015) **77**, 1213-1236. DOI: 10.1007/s11538-015-0083-7.

**Best Student Paper (shared prize): **H. C. Warsinske, S. L. Ashley, J. J. Linderman, B. B. Moore and D. E. Kirschner. Identifying mechanisms of homeostatic signaling in fibroblast differentiation.* Bull Math Biol *(2015) **77**, 1556-1582. DOI: 10.1007/s11538-015-0096-2.

**2015**

**Best Paper: **S.R. McDougall, M.G. Watson, A.H. Devlin, C.A. Mitchell and M.A.J. Chaplain, A hybrid discrete-continuum mathematical model of pattern prediction.* Bull Math Biol *(2012) **74**, 2272-2314. DOI: 10.1007/s11538-012-9754-9.

**Best Student Paper: **S. O’Malley and M. A. Bees, The orientation of swimming bi-flagellates in shear flows.* Bull Math Biol *(2012) **74**, 232-255. DOI: 10.1007/s11538-011-9673-1.

**2013**

**Best Paper: **R. Peña-Miller, D. Lähnemann, H. Schulenburg, M. Ackermann, and R. Beardmore, Selecting against antibiotic-resistant pathogens: optimal treatments in the presence of commensal bacteria.* Bull Math Biol* (2012) **74**, 908-934. DOI: 10.1007/s11538-011-9698-5.

**Best Student Paper: **S.M. Moore, C.A. Manore, V.A. Bokil, E.T. Borer and P.R. Hosseini, Spatiotemporal model of barley and cereal yellow dwarf virus transmission dynamics with seasonality and plant competition.* Bull Math Biol* (2011) **73**, 2707-2730. DOI: 10.1007/s11538-011-9654-4.

**2011**

**Best Research Paper: **W. B. Lindquist and I. D. Chase, Data-based analysis of winner-loser models of hierarchy formation in animals.* Bull Math Biol *(2009) **71**, 556-584. DOI: 10.1007/s11538-008-9371-9.

**Best Educational Paper: **B. R. Kohler, R. J. Swank, J. W. Haefner and J. A. Powell, Leading students to investigate diffusion as a model of brine shrimp movement.* Bull Math Biol *(2010) **72**, 230-257. DOI: 10.1007/s11538-009-9444-4.

**Best Student Paper: **B. Boldin, Persistence and spread of gastro-intestinal infections: the case of enterotoxigenic Escherichia coli in piglets.* Bull Math Biol* (2008) **70**, 2077-2101. DOI: 10.1007/s11538-008-9348-8.

**2009**

**Best Paper: **T. de-Camino-Beck and M. A. Lewis, A new method for calculating net reproductive rate from graph reduction with applications to the control of invasive species.* Bull Math Biol* (2007) **69**, 1341-1354. DOI: 10.1007/s11538-006-9162-0.

**Best Student Paper: **E. Y. Jin and C. M. Reidys, Asymptotic enumeration of RNA structures with pseudoknots.* Bull Math Biol *(2008) **70**, 951-970. DOI: 10.1007/s11538-007-9265-2.